Faculty of Biological Sciences

Prof Bill Kunin

AB, Princeton; MPP, Harvard (Kennedy School); PhD 1991, Univ. Washington (Zoology).
Professor of Ecology
School of Biology

Background: Bill Kunin graduated in Biology from Princeton, received a Masters Degree in Public Policy at Harvard, and a Ph.D. in Zoology at the University of Washington in Seattle. He moved to Britain to take a postdoctoral position at the Centre for Population Biology at Imperial College, and moved to Leeds to take up a lectureship in ecology in 1996. He was promoted to become Reader in spatial ecology in 2006. He lives in a pink house in the suburbs with his wife, Helen Lewis, and their two sets of twins, Nat, Miriam, Toby and Sam. His research focuses on spatial aspects of the interactions between plants and the pollinator and herbivores that feed upon them, but extends to cover aspects of conservation biology, community ecology and biogeographic issues. He manages the BSc programme in Ecology & Environmental Biology, is director of BIOCONS (the European Centre for Biodiversity and Conservation Research) - a Marie Curie Early Stage Training site, and is active in organising the University's interdisciplinary Earth and Biosphere Institute, and UKPopNet - a NERC-funded distributed centre of population biology research. His current research includes several interdisciplinary collaborations, including a NERC PgP project on the range limits of a sub-arctic/alpine plant, two RELU projects on scale effects in farmland biodiversity and sustainable uplands, and coordinating field based research in the EU FP6 ALARM project.

Contact:  Miall 9.19, +44(0) 113 34 32857, email address for  

Research Interests

Spatial aspects of population and community ecology and conservation biology

I am interested in rare species and the factors that influence their populations: their interactions with their environment and with the species that compete with them and feed upon them.  I focus particularly on plant populations, and on their interactions with insect herbivores and pollinators.  Much of my work has relied on manipulative field experiments in which I have set up controlled populations of plants and investigated the effect on pollination and herbivory.  These experiments have highlighted strong positive effects of population density on pollination, while effects on herbivores varies greatly, raising interesting questions for future research.

Natural populations tend to be spatially complex, with individuals patchily distributed across a wide range of spatial scales.  One aspect of my recent work has been the application of fractal and other multi-scale models to describe and predict the spatial and temporal dynamics of such populations.  Modelling work suggests that autocorrelation in dynamics and disturbance may be key in predicting the probability of extinction.   More generally, I have begun work on species distributions, and in particular of the dynamics of populations at the margins of a species’ range.  Some of this recent work has included collaborations with molecular geneticists to examine the factors limiting populations in these marginal areas. 

A third key research area is biodiversity: its maintenance and conservation.  I have conducted research on the role of dispersal across environmental boundaries on diversity (spatial mass effects), one of a range of mechanisms which may help maintain diverse natural communities.  We will soon begin work on the role of surrounding landscapes in explaining local biodiversity in agricultural and upland landscapes, in the UK and across Europe.

My research has been funded by NERC (including a large collaborative Postgenomics project), the EU (as part of the ALARM consortium), the US Forest Service and the interdisciplinary Rural Economy and Land Use and Insect Pollinator Initiative programmes.

Bill Kunin


Current Projects


Faculty Research and Innovation

Studentship information

Undergraduate project topics:

  • Population, community and behavioural ecology; specific topics as for research interests.

Postgraduate studentship areas:

  • The effects of spatial scale on species incidence and interactions

See also:

Modules managed

BLGY2313 - Mediterranean Ecology Field Course
BLGY5102M - Community Ecology
BLGY5156M - Masters Mediterranean Ecology Field Course

Modules taught

BLGY1124/1128 - The Diversity of Life/Living Planet
BLGY1125 - Biology Practicals and Data Analysis
BLGY1128 - Living Planet
BLGY1303 - Tutorials for Biology and Genetics
BLGY1304 - Research Experience and Skills Level 1
BLGY2100 - Enhanced Study Skills for Biologists
BLGY2301 - Research Experience and Skills Level 2
BLGY2313/5156M - Mediterranean Ecology Field Course
BLGY3133 - Advanced Topics in Ecology
BLGY3340 - Biology Research Projects
BLGY5102M - Community Ecology
BLGY5191M - Biodiversity and Conservation MSc and MRes Summer Project
BLGY5198M - Biodiversity and Conservation MRes Research Project 1

Centre membership: The Earth and Biosphere Institute

Group Leader Prof Bill Kunin  (Professor of Ecology)

Spatial aspects of population and community ecology and conservation biology 

Prof Les Firbank  (Professor of Sustainable Agriculture)

Sustainable land management 

Dr Mark Goddard  (Visiting Researcher)

Biodiversity conservation in urban environments 

Dr Richard Gunton  (Research Fellow)

Spatial community ecology 


Andrea Barden (Primary supervisor) 50% FTE
David Brooks (Primary supervisor) 50% FTE
Jordan Chetcuti (Primary supervisor) 100% FTE
Thomas Dally (Primary supervisor) 80% FTE
Johan Fruh (Primary supervisor) 50% FTE
Luca Budello (Co-supervisor) 30% FTE
Solène Guenat (Co-supervisor) 33% FTE
James Hunter (Co-supervisor) 20% FTE
Jens Van Eeckhoven (Co-supervisor) 25% FTE

Carvalheiro LG, Kunin WE, Keil P, Aguirre-Gutierrez J, Ellis WN, Fox R, Groom Q, Hennekens S, Van Landuyt W, Maes D, Van de Meutter F, Michez D, Rasmont P, Ode B, Potts SG, Reemer M, Roberts SPM, Schaminee J, WallisDeVries MF, Biesmeijer JC Species richness declines and biotic homogenisation have slowed down for NW-European pollinators and plants ECOLOGY LETTERS 16 870-878, 2013

Vergeer P, Kunin WE Adaptation at range margins: Common garden trials and the performance of Arabidopsis lyrata across its northwestern European range New Phytologist 197 989-1001, 2013
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Azaele S, Cornell SJ, Kunin WE Downscaling species occupancy from coarse spatial scales Ecological Applications 22 1004-1014, 2012
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Sizling AL, Kunin WE, Sizlingová E, Reif J, Storch D Between geometry and biology: the problem of universality of the species-area relationship. Am Nat 178 602-611, 2011
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Pandit MK, Pocock MJO, Kunin WE Ploidy influences rarity and invasiveness in plants J ECOL 99 1108-1115, 2011

Keil P, Biesmeijer JC, Barendregt A, Reemer M, Kunin WE Biodiversity change is scale-dependent: an example from Dutch and UK hoverflies (Diptera, Syrphidae) ECOGRAPHY 34 392-401, 2011

Polce C, Kunin WE, Biesmeijer JC, Dauber J, Phillips OL, ALARM Field Site Network Alien and native plants show contrasting responses to climate and land use in Europe GLOBAL ECOL BIOGEOGR 20 367-379, 2011

Hodgson JA, Kunin WE, Thomas CD, Benton TG, Gabriel D Comparing organic farming and land sparing: optimizing yield and butterfly populations at a landscape scale ECOL LETT 13 1358-1367, 2010

Gunton RM, Kunin WE Density-dependence at multiple scales in experimental and natural plant populations J ECOL 97 567-580, 2009

Gunton RM, Kunin WE Density effects at multiple scales in an experimental plant population J ECOL 95 435-445, 2007

Senapathi D, Goddard MA, Kunin WE, Baldock KCR Landscape impacts on pollinator communities in temperate systems: evidence and knowledge gaps Functional Ecology 31 26-37, 2017
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Dainese M, Isaac NJ, Powney GD, Bommarco R,Öckinger E, Kuussaari M, Pöyry J, Benton TG, Gabriel D, Hodgson JA, Kunin WE, Lindborg R, Sait SM, Marini L Landscape simplification weakens the association between terrestrial producer and consumer diversity in Europe. Glob Chang Biol -, 2016
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Danino M, Shnerb NM, Azaele S, Kunin WE, Kessler DA The effect of environmental stochasticity on species richness in neutral communities Journal of Theoretical Biology 409 155-164, 2016
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Sizling AL, Pokorny P, Jurickova L, Horackova J, Abraham V, Sizlingova E, Lozek V, Tjorve E, Tjorve KMC, Kunin W Can people change the ecological rules that appear general across space? Global Ecology and Biogeography 25 1072-1084, 2016
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Hicks DM, Ouvrard P, Baldock KCR, Baude M, Goddard MA, Kunin WE, Mitschunas N, Memmott J, Morse H, Nikolitsi M, Osgathorpe LM, Potts SG, Robertson KM, Scott AV, Sinclair F, Westbury DB, Stone GN Food for pollinators: Quantifying the nectar and pollen resources of urban flower meadows PLoS ONE 11 -, 2016
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Socolar JB, Gilroy JJ, Kunin WE, Edwards DP Sparse Data Necessitate Explicit Treatment of Beta-Diversity: A Reply to Bush et al. Trends Ecol Evol 31 338-339, 2016

Baude M, Kunin WE, Boatman ND, Conyers S, Davies N, Gillespie MAK, Morton RD, Smart SM, Memmott J Historical nectar assessment reveals the fall and rise of floral resources in Britain Nature 530 85-88, 2016
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Socolar JB, Gilroy JJ, Kunin WE, Edwards DP How Should Beta-Diversity Inform Biodiversity Conservation? Trends in Ecology and Evolution 31 67-80, 2016
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Šizling AL, Šizlingová E, Tjørve E, Tjørve KMC, Kunin WE How to allow SAR collapse across local and continental scales: A resolution of the controversy between Storch et al. (2012) and Lazarina et al. (2013) Ecography -, 2016
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Ouvrard P, Hicks DM, Mouland M, Nicholls JA, Baldock KCR, Goddard MA, Kunin WE, Potts SG, Thieme T, Veromann E, Stone GN Molecular taxonomic analysis of the plant associations of adult pollen beetles (Nitidulidae: Meligethinae), and the population structure of Brassicogethes aeneus Genome 59 1101-1116, 2016
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Gunton RM, Marsh CJ, Moulherat S, Malchow AK, Bocedi G, Klenke RA, Kunin WE Multicriterion trade-offs and synergies for spatial conservation planning Journal of Applied Ecology -, 2016
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Peay S, Dunn AM, Kunin WE, Mckimm R, Harrod C A method test of the use of electric shock treatment to control invasive signal crayfish in streams Aquatic Conservation: Marine and Freshwater Ecosystems 25 874-880, 2015
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Barwell LJ, Isaac NJB, Kunin WE Measuringβ-diversity with species abundance data Journal of Animal Ecology 84 1112-1122, 2015

Gunton RM, Polce C, Kunin WE Predicting ground temperatures across European landscapes METHODS IN ECOLOGY AND EVOLUTION 6 532-542, 2015

Senapathi D, Carvalheiro LG, Biesmeijer JC, Dodson CA, Evans RL, McKerchar M, Morton RD, Moss ED, Roberts SP, Kunin WE, Potts SG The impact of over 80 years of land cover changes on bee and wasp pollinator communities in England. Proceedings of the Royal Society B: Biological Sciences 282 -, 2015
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Baldock KC, Goddard MA, Hicks DM, Kunin WE, Mitschunas N, Osgathorpe LM, Potts SG, Robertson KM, Scott AV, Stone GN, Vaughan IP, Memmott J Where is the UK's pollinator biodiversity? The importance of urban areas for flower-visiting insects. Proc Biol Sci 282 20142849-, 2015
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Azaele S, Maritan A, Cornell SJ, Suweis S, Banavar JR, Gabriel D, Kunin WE Towards a unified descriptive theory for spatial ecology: Predicting biodiversity patterns across spatial scales Methods in Ecology and Evolution 6 324-332, 2015
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Inclán DJ, Cerretti P, Gabriel D, Benton TG, Sait SM, Kunin WE, Gillespie MAK, Marini L Organic farming enhances parasitoid diversity at the local and landscape scales Journal of Applied Ecology 52 1102-1109, 2015
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Carvalheiro LG, Biesmeijer JC, Benadi G, Fründ J, Stang M, Bartomeus I, Kaiser-Bunbury CN, Baude M, Gomes SI, Merckx V, Baldock KC, Bennett AT, Boada R, Bommarco R, Cartar R, Chacoff N, Dänhardt J, Dicks LV, Dormann CF, Ekroos J, Henson KS, Holzschuh A, Junker RR, Lopezaraiza-Mikel M, Memmott J, Montero-Castaño A, Nelson IL, Petanidou T,Power EF, Rundlöf M, Smith HG, Stout JC, Temitope K, Tscharntke T, Tscheulin T, Vilà M, Kunin WE The potential for indirect effects between co-flowering plants via shared pollinators depends on resource abundance, accessibility and relatedness. Ecol Lett 17 1389-1399, 2014
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Barwell LJ, Azaele S, Kunin WE, Isaac NJB Can coarse-grain patterns in insect atlas data predict local occupancy? DIVERSITY AND DISTRIBUTIONS 20 895-907, 2014

Elbgami T, Kunin WE, Hughes WOH, Biesmeijer JC The effect of proximity to a honeybee apiary on bumblebee colony fitness, development, and performance APIDOLOGIE 45 504-513, 2014

Comont RF, Purse BV, Phillips W, Kunin WE, Hanson M, Lewis OT, Harrington R, Shortall CR, Rondoni G, Roy HE Escape from parasitism by the invasive alien ladybird, Harmonia axyridis INSECT CONSERVATION AND DIVERSITY 7 334-342, 2014

Hoffmann A, Penner J, Vohland K, Cramer W, Doubleday R, Henle K, Kõljalg U, Kühn I, Kunin WE, Negro JJ, Penev L, Rodríguez C, Saarenmaa H, Schmeller DS, Stoev P, Sutherland WJ, Tuama É, Wetzel FT, Häuser CL Improved access to integrated biodiversity data for science, practice, and policy - The European Biodiversity Observation Network (EU BON) Nature Conservation 6 49-65, 2014
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Spracklen BD, Lane JV, Kunin WE, Spracklen DV, Williams N Regeneration of native broadleaved species on clearfelled conifer plantations in upland Britain Forest Ecology and Management 310 204-212, 2013
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Shackelford G, Steward PR, Benton TG, Kunin WE, Potts SG, Biesmeijer JC, Sait SM Comparison of pollinators and natural enemies: a meta-analysis of landscape and local effects on abundance and richness in crops. Biol Rev Camb Philos Soc 88 1002-1021, 2013
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Carvalheiro LG, Kunin WE, Biesmeijer JC Corrigendum to Carvalheiro et al. [Ecol. Lett., 16, (2013) 870-878] Ecology Letters 16 1416-1417, 2013

Kunin WE No bias behind pollinator research NATURE 502 303-303, 2013

Dicks LV, Abrahams A, Atkinson J, Biesmeijer J, Bourn N, Brown C, Brown MJF, Carvell C, Connolly C, Cresswell JE, Croft P, Darvill B, De Zylva P, Effingham P, Fountain M, Goggin A, Harding D, Harding T, Hartfield C, Heard MS, Heathcote R, Heaver D, Holland J, Howe M, Hughes B, Huxley T, Kunin WE, Little J, Mason C, Memmott J, Osborne J, Pankhurst T, Paxton RJ, Pocock MJO, Potts SG, Power EF, Raine NE, Ranelagh E, Roberts S, Saunders R, Smith K, Smith RM, Sutton P, Tilley LAN, Tinsley A, Tonhasca A, Vanbergen AJ, Webster S, Wilson A, Sutherland WJ Identifying key knowledge needs for evidence-based conservation of wild insect pollinators: a collaborative cross-sectoral exercise INSECT CONSERVATION AND DIVERSITY 6 435-446, 2013

Gabriel D, Sait SM, Kunin WE, Benton TG Food production vs. biodiversity: comparing organic and conventional agriculture JOURNAL OF APPLIED ECOLOGY 50 355-364, 2013

Sizling AL, Kunin WE, Storch D Taxon-and-Area Invariances, Maximum Entropy, and the Species-Area Relationship. Am Nat 181 288-290, 2013

Bradter U, Kunin WE, Altringham JD, Thom TJ, Benton TG Identifying appropriate spatial scales of predictors in species distribution models with the random forest algorithm Methods in Ecology and Evolution 4 167-174, 2013
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Moustakas A, Kunin WE, Cameron TC, Sankaran M Facilitation or competition? Tree effects on grass biomass across a precipitation gradient. PLoS One 8 e57025-, 2013
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Reed MS, Hubacek K, Bonn A, Burt TP, Holden J, Stringer LC, Beharry-Borg N, Buckmaster S, Chapman D, Chapman PJ, Clay GD, Cornell SJ, Dougill AJ, Evely AC, Fraser EDG, Jin N, Irvine BJ, Kirkby MJ, Kunin WE, Prell C, Quinn CH, Slee B, Stagl S, Termansen M, Thorp S, Worrall F Anticipating and managing future trade-offs and complementarities between ecosystem services Ecology and Society 18 -, 2013
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Carvalheiro LG, Kunin WE, Keil P, Aguirre-Gutiérrez J, Ellis WN, Fox R, Groom Q, Hennekens S, Van Landuyt W, Maes D, Van de Meutter F, Michez D, Rasmont P, Ode B, Potts SG, Reemer M, Roberts SPM, Schaminée J, Wallisdevries MF, Biesmeijer JC Species richness declines and biotic homogenisation have slowed down for NW-European pollinators and plants Ecology Letters 16 870-878, 2013
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Sutherland WJ, Freckleton RP, Godfray HCJ, Beissinger SR, Benton T, Cameron DD, Carmel Y, Coomes DA, Coulson T, Emmerson MC, Hails RS, Hays GC, Hodgson DJ, Hutchings MJ, Johnson D, Jones JPG, Keeling MJ, Kokko H, Kunin WE, Lambin X, Lewis OT, Malhi Y, Mieszkowska N, Milner-Gulland EJ, Norris K, Phillimore AB, Purves DW, Reid JM, Reuman DC, Thompson K, Travis JMJ, Turnbull LA, Wardle DA, Wiegand T Identification of 100 fundamental ecological questions Journal of Ecology 101 58-67, 2013
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Gillingham PK, Huntley B, Kunin WE, Thomas CD The effect of spatial resolution on projected responses to climate warming DIVERSITY AND DISTRIBUTIONS 18 990-1000, 2012

Gillingham PK, Palmer SCF, Huntley B, Kunin WE, Chipperfield JD, Thomas CD The relative importance of climate and habitat in determining the distributions of species at different spatial scales: a case study with ground beetles in Great Britain ECOGRAPHY 35 831-838, 2012

Keil P, Schweiger O, Kuehn I, Kunin WE, Kuussaari M, Settele J, Henle K, Brotons L, Pe'er G, Lengyel S, Moustakas A, Steinicke H, Storch D Patterns of beta diversity in Europe: the role of climate, land cover and distance across scales JOURNAL OF BIOGEOGRAPHY 39 1473-1486, 2012

Sutherland LA, Gabriel D, Hathaway-Jenkins L, Pascual U, Schmutz U, Rigby D, Godwin R, Sait SM, Sakrabani R, Kunin WE, Benton TG, Stagl S The 'Neighbourhood Effect': A multidisciplinary assessment of the case for farmer co-ordination in agri-environmental programmes Land Use Policy 29 502-512, 2012
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Segurado P, Kunin WE, Filipe AF, Araujo MB Patterns of coexistence of two species of freshwater turtles are affected by spatial scale BASIC AND APPLIED ECOLOGY 13 371-379, 2012

Nielsen A, Dauber J, Lamborn E, Tscheulin T, Vaitis M, Vivarelli D, Petanidou T, Nielsen A, Nielsen A, Reitan T, Dauber J, Kunin WE, Biesmeijer JC, Dauber J, Stout JC, Lamborn E, Potts SG, Roberts S, Tscheulin T, Jauker B, Jauker B, Moora M, Sõber V, Settele J, Steffan-Dewenter I, Vivarelli D Pollinator community responses to the spatial population structure of wild plants: A pan-European approach Basic and Applied Ecology -, 2012
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Nielsen A, Steffan-Dewenter I, Westphal C, Messinger O, Potts SG, Roberts SPM, Settele J, Szentgyoergyi H, Vaissiere BE, Vaitis M, Woyciechowski M, Bazos I, Biesmeijer JC, Bommarco R, Kunin WE, Tscheulin T, Lamborn E, Petanidou T Assessing bee species richness in two Mediterranean communities: importance of habitat type and sampling techniques ECOLOGICAL RESEARCH 26 969-983, 2011

Bradter U, Thom TJ, Altringham JD, Kunin WE, Benton TG Prediction of National Vegetation Classification communities in the British uplands using environmental data at multiple spatial scales, aerial images and the classifier random forest J APPL ECOL 48 1057-1065, 2011

Vergeer P, Kunin WE Life history variation in Arabidopsis lyrata across its range: effects of climate, population size and herbivory OIKOS 120 979-990, 2011

Moora M, Berger S, Davison J, Opik M, Bommarco R, Bruelheide H, Kuhn I, Kunin WE, Metsis M, Rortais A, Vanatoa A, Vanatoa E, Stout JC, Truusa M, Westphal C, Zobel M, Walther GR Alien plants associate with widespread generalist arbuscular mycorrhizal fungal taxa: evidence from a continental-scale study using massively parallel 454 sequencing J BIOGEOGR 38 1305-1317, 2011

Pysek P, Jarosik V, Chytry M, Danihelka J, Kuhn I, Pergl J, Tichy L, Biesmeijer JC, Ellis WN, Kunin WE, Settele J Successful invaders co-opt pollinators of native flora and accumulate insect pollinators with increasing residence time ECOL MONOGR 81 277-293, 2011

Thomas CD, Hill JK, Anderson BJ, Bailey S, Beale CM, Bradbury RB, Bulman CR, Crick HQP, Eigenbrod F, Griffiths HM, Kunin WE, Oliver TH, Walmsley CA, Watts K, Worsfold NT, Yardley T A framework for assessing threats and benefits to species responding to climate change METHODS ECOL EVOL 2 125-142, 2011

Potts SG, Biesmeijer JC, Bommarco R, Felicioli A, Fischer M, Jokinen P, Kleijn D, Klein AM, Kunin WE, Neumann P, Penev LD, Petanidou T, Rasmont P, Roberts SPM, Smith HG, Sorensen PB, Steffan-Dewenter I, Vaissiere BE, Vila M, Vujic A, Woyciechowski M, Zobel M, Settele J, Schweiger O Developing European conservation and mitigation tools for pollination services: approaches of the STEP (Status and Trends of European Pollinators) project J APICULT RES 50 152-164, 2011

Suggitt AJ, Gillingham PK, Hill JK, Huntley B, Kunin WE, Roy DB, Thomas CD Habitat microclimates drive fine-scale variation in extreme temperatures OIKOS 120 1-8, 2011

Marini L, Bona E, Kunin WE, Gaston KJ Exploring anthropogenic and natural processes shaping fern species richness along elevational gradients J BIOGEOGR 38 78-88, 2011

Hooper EJ, Duckett JG, Cuming AC, Kunin WE, Pressel S Ephemerella readeri Mull. Hal. (Physcomitrella readeri (Mull. Hal.) I.G. Stone&G.A.M. Scott, Funariidae, Bryophyta): a genus and species new to Europe J BRYOL 32 256-264, 2010

Gabriel D, Sait SM, Hodgson JA, Schmutz U, Kunin WE, Benton TG Scale matters: the impact of organic farming on biodiversity at different spatial scales ECOL LETT 13 858-869, 2010

Potts SG, Biesmeijer JC, Kremen C, Neumann P, Schweiger O, Kunin WE Global pollinator declines: trends, impacts and drivers TRENDS ECOL EVOL 25 345-353, 2010

Hammen VC, Biesmeijer JC, Bommarco R, Budrys E, Christensen TR, Fronzek S, Grabaum R, Jaksic P, Klotz S, Kramarz P, Kroel-Dulay G, Kuhn I, Mirtl M, Moora M, Petanidou T, Pino J, Potts SG, Rortais A, Schulze CH, Steffan-Dewenter I, Stout J, Szentgyorgyi H, Vighi M, Vujic A, Westphal C, Wolf T, Zavala G, Zobel M, Settele J, Kunin WE Establishment of a cross-European field site network in the ALARM project for assessing large-scale changes in biodiversity ENVIRON MONIT ASSESS 164 337-348, 2010

Dauber J, Biesmeijer JC, Gabriel D, Kunin WE, Lamborn E, Meyer B, Nielsen A, Potts SG, Roberts SPM, Sober V, Settele J, Steffan-Dewenter I, Stout JC, Teder T, Tscheulin T, Vivarelli D, Petanidou T Effects of patch size and density on flower visitation and seed set of wild plants: a pan-European approach J ECOL 98 188-196, 2010

Chapman DS, Bonn A, Kunin WE, Cornell SJ Random Forest characterization of upland vegetation and management burning from aerial imagery J BIOGEOGR 37 37-46, 2010

Reed MS, Bonn A, Slee W, Beharry-Borg N, Birch J, Brown I, Burt TP, Chapman D, Chapman PJ, Clay GD, Cornell SJ, Fraser EDG, Glass JH, Holden J, Hodgson JA, Hubacek K, Irvine B, Jin N, Kirkby MJ, Kunin WE, Moore O, Moseley D, Prell C, Price MF, Quinn CH, Redpath S, Reid C, Stagl S, Stringer LC, Termansen M, Thorp S, Towers W, Worrall F The future of the uplands LAND USE POLICY 26 S204-S216, 2009

Kunin WE, Vergeer P, Kenta T, Davey MP, Burke T, Woodward FI, Quick P, Mannarelli ME, Watson-Haigh NS, Butlin R Variation at range margins across multiple spatial scales: environmental temperature, population genetics and metabolomic phenotype P R SOC B 276 1495-1506, 2009

Chapman DS, Termansen M, Quinn CH, Jin NL, Bonn A, Cornell SJ, Fraser EDG, Hubacek K, Kunin WE, Reed MS Modelling the coupled dynamics of moorland management and upland vegetation J APPL ECOL 46 278-288, 2009

Gabriel D, Carver SJ, Durham H, Kunin WE, Palmer RC, Sait SM, Stagl S, Benton TG The spatial aggregation of organic farming in England and its underlying environmental correlates J APPL ECOL 46 323-333, 2009

Chapman DS, Cornell SJ, Kunin WE Interactions between harvesting, noise and territoriality in a model of red grouse population cycles J ANIM ECOL 78 476-484, 2009

Tjorve E, Kunin WE, Polce C, Tjorve KMC Species-area relationship: separating the effects of species abundance and spatial distribution J ECOL 96 1141-1151, 2008

Westphal C, Bommarco R, Carre G, Lamborn E, Morison N, Petanidou T, Potts SG, Roberts SPM, Szentgyorgyi H, Tscheulin T, Vaissiere BE, Woyciechowski M, Biesmeijer JC, Kunin WE, Settele J, Steffan-Dewenter I MEASURING BEE DIVERSITY IN DIFFERENT EUROPEAN HABITATS AND BIOGEOGRAPHICAL REGIONS ECOL MONOGR 78 653-671, 2008

Kunin WE On comparative analyses involving non-heritable traits: why half a loaf is sometimes worse than none EVOL ECOL RES 10 787-796, 2008

Vergeer P, van den Berg LLJ, Bulling MT, Ashmore MR, Kunin WE Geographical variation in the response to nitrogen deposition in Arabidopsis lyrata petraea NEW PHYTOL 179 129-141, 2008

Fyllas NM, Phillips OL, Kunin WE, Matsinos YG, Troumbis AI Development and parameterization of a general forest gap dynamics simulator for the North-eastern Mediterranean Basin (GREek FOrest Species) ECOL MODEL 204 439-456, 2007

Gaston KJ, Charman K, Jackson SF, Armsworth PR, Bonn A, Briers RA, Callaghan CSQ, Catchpole R, Hopkins J, Kunin WE, Latham J, Opdam P, Stoneman R, Stroud DA, Tratt R The ecological effectiveness of protected areas: The United Kingdom BIOL CONSERV 132 76-87, 2006

Van Der Velden NK, Kunin WE, Carver SJ Piecewise linear regression models to characterize understorey edge extents across a clearcut-forest transition Ecological Applications 0-39, 2006
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Sutherland WJ, Armstrong-Brown S, Armsworth PR, Brereton T, Brickland J, Campbell CD, Chamberlain DE, Cooke AI, Dulvy NK, Dusic NR, Fitton M, Freckleton RP, Godfray HCJ, Grout N, Harvey HJ, Hedley C, Hopkins JJ, Kift NB, Kirby J, Kunin WE, Macdonald DW, Marker B, Naura M, Neale AR, Oliver T, Osborn D, Pullin AS, Shardlow MEA, Showler DA, Smith PL, Smithers RJ, Solandt JL, Spencer J, Spray CJ, Thomas CD, Thompson J, Webb SE, Yalden DW, Watkinson AR The identification of 100 ecological questions of high policy relevance in the UK J APPL ECOL 43 617-627, 2006

Biesmeijer JC, Roberts SPM, Reemer M, Ohlemuller R, Edwards M, Peeters T, Schaffers AP, Potts SG, Kleukers R, Thomas CD, Settele J, Kunin WE Parallel declines in pollinators and insect-pollinated plants in Britain and the Netherlands SCIENCE 313 351-354, 2006

Metcalfe DB, Kunin WE The effects of plant density upon pollination success, reproductive effort and fruit parasitism in Cistus ladanifer L. (Cistaceae) PLANT ECOL 185 41-47, 2006

Segurado P, Araujo MB, Kunin WE Consequences of spatial autocorrelation for niche-based models Journal of Applied Ecology 43 433-444, 2006
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Pocock MJO, Hartley S, Telfer MG, Preston CD, Kunin WE Ecological correlates of range structure in rare and scarce British plants J ECOL 94 581-596, 2006

Kallimanis AS, Kunin WE, Halley JM, Sgardelis SP Patchy disturbance favours longer dispersal distance EVOL ECOL RES 8 529-541, 2006

Kallimanis AS, Kunin WE, Halley JM, Sgardelis SP Metapopulation extinction risk under spatially autocorrelated disturbance CONSERV BIOL 19 534-546, 2005

Wilson RJ, Thomas CD, Fox R, Roy DB, Kunin WE Spatial patterns in species distributions reveal biodiversity change NATURE 432 393-396, 2004

Corne SA, Carver SJ, Kunin WE, Lennon JJ, Van Hees WWS Predicting Forest Attributes in Southeast Alaska Using Artificial Neural Networks Forest Science 50 259-276, 2004
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Halley JM, Hartley S, Kallimanis AS, Kunin WE, Lennon JJ, Sgardelis SP Uses and abuses of fractal methodology in ecology Ecology Letters 7 254-271, 2004
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Hartley S, Kunin WE Scale dependency of rarity, extinction risk, and conservation priority CONSERV BIOL 17 1559-1570, 2003

Hartley S, Kunin WE, Lennon J, Pocock MJO Coherence and discontinuity in the scaling of species' distribution patterns Proceedings of the Royal Society of London. Series B. Biological Sciences 271 81-88, 2003
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Lennon JJ, Kunin WE, Hartley S Fractal species distributions do not produce power-law species-area relationships OIKOS 97 378-386, 2002

Lennon J, Kunin WE, Corne SA, Carver SJ, Van Hees WWS Are Alaskan trees found in locally more favourable sites in marginal areas? Global Ecology and Biogeography Letters 11 103-114, 2002
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Kunin WE, Hartley S, Lennon J Scaling down: on the challenge of estimating abundance from occurance patterns The American Naturalist 156 560-566, 2000

Halley JM, Kunin WE Extinction risk and the 1/f-family of noise models Theoretical Population Biology 56 215-230, 1999

Thomas CD, Kunin WE The Spatial structure of populations Journal of Animal Ecology 68 647-657, 1999

Kunin WE Patterns of herbivore incidence on experimental arrays and field populations of ragwort (Senecio jacobaea) Oikos 84 515-525, 1999

Kunin WE Extrapolating species abundance across spatial scales Science 281 1513-1515, 1998
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Kunin WE Biodiversity at the Edge: a Test of the Importance of Spatial 'Mass Effects' in the Rothamsted Park Grass Experiments Proceedings of the National Academy of Sciences of the United States of America 95 207-212, 1998
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Kunin WE Population size and density effects in pollination: pollinator foraging and plant reproductive success in experimental arrays of Brassica kaber Journal of Ecology 85 225-234, 1997
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Kunin WE, Shmida A Plant reproductive traits as a function of local, regional, and global abundance Conservation Biology 11 183-192, 1997
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Kunin WE Sample shapes, spatial scale and species counts: implications for reserve design Biological Conservation 82 369-377, 1997

Kunin WE Pollinator foraging strategies in mixed floral arrays: density effects and floral constancy Theoretical Population Biology 49 232-263, 1996

Kunin WE Towards an asymmetric index of community similarity Oikos 73 442-446., 1995

Kunin WE Density dependent foraging in the harvester ant Messor ebeninus: two experiments Oecologia 98 328-335., 1994

Kunin WE The biology of rarity: patterns, causes, and consequences. Trends in Ecology and Evolution 8 298-301., 1993

Kunin WE Sex and the single mustard: population density and pollinator behaviour effects on seed-set. Ecology 74 2145-2160., 1993

Kunin WE, Lennon J Spatial scale and species diversity: building species-area curves from species incidence In Dryland Biodiversity ,

Gavish Y, Marsh CJ, Kuemmerlen M, Stoll S, Haase P, Kunin WE Accounting for biotic interactions through alpha-diversity constraints in stacked species distribution models Methods in Ecology and Evolution -,
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