Prof Keith Hamer

BSc, MSc, Manchester; PhD 1990, Glasgow.
Professor of Animal Ecology
School of Biology

Contact: Manton 8.05, +44(0) 113 34 32983, email address for  

You can read more about Prof Hamer's interests here:

Research Interests

Population and Behavioural Ecology; Conservation Biology

Gannets on rockMy research focuses on wildlife ecology and the conservation of biodiversity in natural and managed landscapes and seascapes, with a particular emphasis on trophic ecology – what animals eat, how they get enough, and how this determines both their responses to environmental change and their contributions to broader ecosystem-level processes. Much of this work involves birds and insects (especially butterflies, dung beetles and ants), mainly in marine, forested and agricultural ecosystems. Current  topics include the importance of individual specialization in the foraging strategies of long-distance predators and the role of trophic flexibility in buffering impacts of environmental change. We are also investigating the patterns, causes and consequences of changes in biodiversity following degradation and fragmentation of tropical forests, and examining methods for improving biodiversity in oil palm landscapes of SE Asia and West Africa. 

Current projects:

  • Impacts of offshore windfarms on populations and demography of gannets: reducing uncertainty in risk assessments (NERC iCASE)
  • Enhancing the benefits of tropical peatland restoration for local communities and ecosystem processes (NERC Newton Fund)
  • Promoting biodiversity in sustainable oil-palm landcapes for West African smallholders (DEFRA Darwin Initiative;
  • Individual foraging strategies of gannets: efficiency, effort and success (NERC)
  • Socially and Environmentally Sustainable Oil Palm Research (SEnSOR;
  • Ecology and epidemiology of Trichomonas gallinae in turtle doves (RSPB)
  • Individual consistency and constraint in the foraging behaviour of shags (NERC)  
  • Three-dimensional foraging behaviour of gannets: integrating location, flight height and dive depth (NERC)

Faculty Research and Innovation

Studentship information

Undergraduate project topics:

  • 1. Drivers of distribution patterns: three potential projects examining the importance of different processes and species traits in determining distribution patterns at different spatial scales
  • 2. Impacts of habitat fragmentation: two potential projects using analysis of nestedness to examine impacts of forest fragmentation on birds and to test predictions of island biogeography theory
  • 3. Impacts of climate change on seabirds: a meta-analysis of published data

Postgraduate studentship areas:

  • 1. Foraging strategies, parental care and life history-environment interactions in birds
  • 2. Species richness and ecosystem functioning in natural, managed, degraded, fragmented and converted forests
  • 3. Biodiversity and sustainable management of mosaic landscapes
  • 4. Land-sparing vs land-sharing approaches to sustainable development

See also:

Modules managed

BLGY2144 - Population and Community Ecology
BLGY5118M - Population Dynamics

Modules taught

BLGY1128 - Living Planet
BLGY1304 - Research Experience and Skills Level 1
BLGY2144 - Population and Community Ecology
BLGY2301 - Research Experience and Skills Level 2
BLGY3247 - Advanced Topics in Conservation Science
BLGY3345 - Biology Integrated Research Projects
BLGY3395 - Advanced Research Skills and Experience
BLGY3396 - Research Literature Review
BLGY5118M - Population Dynamics
BLGY5191M - Biodiversity and Conservation MSc and MRes Summer Project

Academic roles

School Director of Research and Innovation - School of Biology


Member of Athena Swan Working Group

Centre membership: The Earth and Biosphere Institute

Group Leader Prof Keith Hamer  (Professor of Animal Ecology)

Population and Behavioural Ecology; Conservation Biology 

Dr Jenny Dunn  (Visiting Research Fellow)

Farmland bird ecology and conservation 


Ruth Jeavons (Primary supervisor) 75% FTE
Jude Lane (Primary supervisor) 90% FTE
Linda Ofosuhene (Primary supervisor) 100% FTE
Christopher Pollock (Primary supervisor) 90% FTE
Michael Sasu (Primary supervisor) 100% FTE
Andrea Barden (Co-supervisor) 50% FTE
Katie Cook (Co-supervisor) 20% FTE
Carrie Easter (Co-supervisor) 30% FTE
Alexander Evans (Co-supervisor) 10% FTE

Qie L, Lewis SL, Sullivan MJP, Lopez-Gonzalez G, Pickavance GC, Sunderland T, Ashton P, Hubau W, Abu Salim K, Aiba S-I, Banin LF, Berry N, Brearley FQ, Burslem DFRP, Dancak M, Davies SJ, Fredriksson G, Hamer KC, Hedl R, Kho LK, Kitayama K, Krisnawati H, Lhota S, Malhi Y, Maycock C, Metali F, Mirmanto E, Nagy L, Nilus R, Ong R, Pendry CA, Poulsen AD, Primack RB, Rutishauser E, Samsoedin I, Saragih B, Sist P, Slik JWF, Sukri RS, Svatek M, Tan S, Tjoa A, van Nieuwstadt M, Vernimmen RRE, Yassir I, Susan Kidd P, Fitriadi M, Ideris NKH, Serudin RM, Lim LSA, Saparudin MS, Phillips OL Long-term carbon sink in Borneo's forests halted by drought and vulnerable to edges (vol 8, 2017) NATURE COMMUNICATIONS 9, 2018

Bennison A, Bearhop S, Bodey TW, Votier SC, Grecian WJ, Wakefield ED, Hamer KC, Jessopp M Search and foraging behaviors from movement data: A comparison of methods Ecology and Evolution 8 13-24, 2018
View abstract

James Grecian W, Lane JV, Michelot T, Wade HM, Hamer KC Understanding the ontogeny of foraging behaviour: Insights from combining marine predator bio-logging with satellite-derived oceanography in hidden Markov models Journal of the Royal Society Interface 15, 2018
View abstract

Qie L, Lewis SL, Sullivan MJP, Lopez-Gonzalez G, Pickavance GC, Sunderland T, Ashton P, Hubau W, Abu Salim K, Aiba SI, Banin LF, Berry N, Brearley FQ, Burslem DFRP, Dančák M, Davies SJ, Fredriksson G, Hamer KC, Hédl R, Kho LK, Kitayama K, Krisnawati H, Lhota S, Malhi Y, Maycock C, Metali F, Mirmanto E, Nagy L, Nilus R, Ong R, Pendry CA, Poulsen AD, Primack RB, Rutishauser E, Samsoedin I, Saragih B, Sist P, Slik JWF, Sukri RS, Svátek M, Tan S, Tjoa A, Van Nieuwstadt M, Vernimmen RRE, Yassir I, Kidd PS, Fitriadi M, Ideris NKH, Serudin RM, Abdullah Lim LS, Saparudin MS, Phillips OL Long-term carbon sink in Borneo's forests halted by drought and vulnerable to edge effects Nature Communications 8, 2017
View abstract

Ross SRPJ, Hassall C, Hoppitt WJE, Edwards FA, Edwards DP, Hamer KC Incorporating intraspecific trait variation into functional diversity: Impacts of selective logging on birds in Borneo Methods in Ecology and Evolution 8 1499-1505, 2017
View abstract

Votier SC, Fayet AL, Bearhop S, Bodey TW, Clark BL, Grecian J, Guilford T, Hamer KC, Jeglinski JWE, Morgan G, Wakefield E, Patrick SC Effects of age and reproductive status on individual foraging site fidelity in a long-lived marine predator. Proceedings of the Royal Society B: Biological Sciences 284, 2017
View abstract

Marx M, Reiner G, Willems H, Rocha G, Hillerich K, Masello JF, Mayr SL, Moussa S, Dunn JC, Thomas RC, Goodman SJ, Hamer KC, Metzger B, Cecere JG, Spina F, Koschkar S, Calderón L, Romeike T, Quillfeldt P High prevalence of Trichomonas gallinae in wild columbids across western and southern Europe Parasites and Vectors 10 1-11, 2017
View abstract

Dunn JC, Stockdale JE, Bradford EL, McCubbin A, Morris AJ, Grice PV, Goodman SJ, Hamer KC High rates of infection by blood parasites during the nestling phase in UK Columbids with notes on ecological associations Parasitology 144 622-628, 2017
View abstract

Dunn JC, Hamer KC, Benton TG Dynamics of phenotypic change: wing length declines in a resident farmland passerine despite survival advantage of longer wings Ibis 159 152-157, 2017
View abstract

Riddick SN, Blackall TD, Dragosits U, Tang YS, Moring A, Daunt F, Wanless S, Hamer KC, Sutton MA High temporal resolution modelling of environmentally-dependent seabird ammonia emissions: Description and testing of the GUANO model Atmospheric Environment 161 48-60, 2017
View abstract

Edwards FA, Finan J, Graham LK, Larsen TH, Wilcove DS, Hsu WW, Chey VK, Hamer KC The impact of logging roads on dung beetle assemblages in a tropical rainforest reserve Biological Conservation 205 85-92, 2017
View abstract

Dunn JC, Stockdale JE, McCubbin A, Thomas RC, Goodman SJ, Grice PV, Morris AJ, Hamer KC, Symondson WOC Non-cultured faecal and gastrointestinal seed samples fail to detect Trichomonad infection in clinically and sub-clinically infected columbid birds Conservation Genetics Resources 8 97-99, 2016
View abstract

Cleasby IR, Wakefield ED, Bearhop S, Bodey TW, Votier SC, Hamer KC Three-dimensional tracking of a wide-ranging marine predator: Flight heights and vulnerability to offshore wind farms Journal of Applied Ecology 52 1474-1482, 2015
View abstract

Wakefield ED, Cleasby IR, Bearhop S, Bodey TW, Davies RD, Miller PI, Newton J, Votier SC, Hamer KC Long-term individual foraging site fidelity-why some gannets don't change their spots Ecology 96 3058-3074, 2015
View abstract

Dunn JC, Hamer KC, Benton TG Anthropogenically-mediated density dependence in a declining farmland bird PLoS ONE 10, 2015
View abstract

Patrick SC, Bearhop S, Bodey TW, Grecian WJ, Hamer KC, Lee J, Votier SC Individual seabirds show consistent foraging strategies in response to predictable fisheries discards Journal of Avian Biology 46 431-440, 2015
View abstract

Hamer KC, Newton RJ, Edwards FA, Benedick S, Bottrell SH, Edwards DP Impacts of selective logging on insectivorous birds in Borneo: The importance of trophic position, body size and foraging height Biological Conservation 188 82-88, 2015
View abstract

Stockdale JE, Dunn JC, Goodman SJ, Morris AJ, Sheehan DK, Grice PV, Hamer KC The protozoan parasite Trichomonas gallinae causes adult and nestling mortality in a declining population of European Turtle Doves, Streptopelia turtur Parasitology 142 490-498, 2015
View abstract

Russell DJF, Wanless S, Collingham YC, Anderson BJ, Beale C, Reid JB, Huntley B, Hamer KC Beyond climate envelopes: bio-climate modelling accords with observed 25-year changes in seabird populations of the British Isles Diversity and Distributions: a journal of conservation biogeography 21 211-222, 2015
View abstract

Cleasby IR, Wakefield ED, Bodey TW, Davies RD, Patrick SC, Newton J, Votier SC, Bearhop S, Hamer KC Sexual segregation in a wide-ranging marine predator is a consequence of habitat selection Marine Ecology Progress Series 518 1-12, 2015
View abstract

Russell DJF, Wanless S, Collingham YC, Huntley B, Hamer KC Predicting future European breeding distributions of British seabird species under climate change and unlimited/no dispersal scenarios Diversity 7 342-359, 2015
View abstract

Edwards DP, Magrach A, Woodcock P, Ji Y, Lim NTL, Edwards FA, Larsen TH, Hsu WW, Benedick S, Khen CV, Chung AYC, Reynolds G, Fisher B, Laurance WF, Wilcove DS, Hamer KC, Yu DW Selective-logging and oil palm: Multitaxon impacts, biodiversity indicators, and trade-offs for conservation planning Ecological Applications 24 2029-2049, 2014
View abstract

Tawatao N, Lucey JM, Senior M, Benedick S, Vun Khen C, Hill JK, Hamer KC Biodiversity of leaf-litter ants in fragmented tropical rainforests of Borneo: the value of publically and privately managed forest fragments Biodiversity and Conservation 23 3113-3126, 2014
View abstract

Bodey TW, Jessopp MJ, Votier SC, Gerritsen HD, Cleasby IR, Hamer KC, Patrick SC, Wakefield ED, Bearhop S Seabird movement reveals the ecological footprint of fishing vessels. Current Biology 24 R514-R515, 2014
View abstract

Dunn JC, Goodman SJ, Benton TG, Hamer KC Active blood parasite infection is not limited to the breeding season in a declining farmland bird Journal of Parasitology 100 260-266, 2014
View abstract

Edwards FA, Edwards DP, Larsen TH, Hsu WW, Benedick S, Chung A, Khen CV, Wilcove DS, Hamer KC Does logging and forest conversion to oil palm agriculture alter functional diversity in a biodiversity hotspot? ANIMAL CONSERVATION 17 163-173, 2014

Edwards FA, Edwards DP, Sloan S, Hamer KC Sustainable management in crop monocultures: the impact of retaining forest on oil palm yield. PLoS One 9 e91695-, 2014
View abstract

Patrick SC, Bearhop S, Gremillet D, Lescroel A, Grecian WJ, Bodey TW, Hamer KC, Wakefield E, Le Nuz M, Votier SC Individual differences in searching behaviour and spatial foraging consistency in a central place marine predator OIKOS 123 33-40, 2014

Lucey JM, Tawatao N, Senior MJM, Khen CV, Benedick S, Hamer KC, Woodcock P, Newton RJ, Bottrell SH, Hill JK Tropical forest fragments contribute to species richness in adjacent oil palm plantations BIOLOGICAL CONSERVATION 169 268-276, 2014

Edwards DP, Gilroy JJ, Woodcock P, Edwards FA, Larsen TH, Andrews DJR, Derhe MA, Docherty TDS, Hsu WW, Mitchell SL, Ota T, Williams LJ, Laurance WF, Hamer KC, Wilcove DS Land-sharing versus land-sparing logging: reconciling timber extraction with biodiversity conservation GLOBAL CHANGE BIOLOGY 20 183-191, 2014

Thaxter CB, Daunt F, Grémillet D, Harris MP, Benvenuti S, Watanuki Y, Hamer KC, Wanless S Modelling the effects of prey size and distribution on prey capture rates of two sympatric marine predators PLoS ONE 8, 2013
View abstract

Ji Y, Ashton L, Pedley SM, Edwards DP, Tang Y, Nakamura A, Kitching R, Dolman PM, Woodcock P, Edwards FA, Larsen TH, Hsu WW, Benedick S, Hamer KC, Wilcove DS, Bruce C, Wang X, Levi T, Lott M, Emerson BC, Yu DW Reliable, verifiable and efficient monitoring of biodiversity via metabarcoding ECOLOGY LETTERS 16 1245-1257, 2013

Edwards DP, Woodcock P, Newton RJ, Edwards FA, Andrews DJR, Docherty TDS, Mitchell SL, Ota T, Benedick S, Bottrell SH, Hamer KC Trophic Flexibility and the Persistence of Understory Birds in Intensively Logged Rainforest Conservation Biology 27 1079-1086, 2013
View abstract

Dunn JC, Goodman SJ, Benton TG, Hamer KC Avian blood parasite infection during the non-breeding season: an overlooked issue in declining populations? BMC Ecology 13, 2013
View abstract

Lennon RJ, Dunn JC, Stockdale JE, Goodman SJ, Morris AJ, Hamer KC Trichomonad parasite infection in four species of Columbidae in the UK. Parasitology 140 1368-1376, 2013
View abstract

Hamer KC Procellariiformes In Reference Module in Earth Systems and Environmental Sciences, 2013
View abstract

Wakefield ED, Bodey TW, Bearhop S, Blackburn J, Colhoun K, Davies R, Dwyer RG, Green J, Gremillet D, Jackson AL, Jessopp MJ, Kane A, Langston RHW, Lescroel A, Murray S, Le Nuz M, Patrick SC, Peron C, Soanes L, Wanless S, Votier SC, Hamer KC Space Partitioning Without Territoriality in Gannets Science 341 68-70, 2013
View abstract

Woodcock P, Edwards DP, Newton RJ, Vun Khen C, Bottrell SH, Hamer KC Impacts of intensive logging on the trophic organisation of ant communities in a biodiversity hotspot. PLoS One 8 e60756-, 2013
View abstract

Edwards FA, Edwards DP, Hamer KC, Davies RG Impacts of logging and conversion of rainforest to oil palm on the functional diversity of birds in Sundaland IBIS 155 313-326, 2013

Davies RD, Wanless S, Lewis S, Hamer KC Density-dependent foraging and colony growth in a pelagic seabird species under varying environmental conditions. Marine Ecology Progress Series 485 287-294, 2013
View abstract

Senior MJM, Hamer KC, Bottrell S, Edwards DP, Fayle TM, Lucey JM, Mayhew PJ, Newton R, Peh KS-H, Sheldon FH, Stewart C, Styring AR, Thom MDF, Woodcock P, Hill JK Trait-dependent declines of species following conversion of rain forest to oil palm plantations BIODIVERSITY AND CONSERVATION 22 253-268, 2013

Fauset S, Baker TR, Lewis SL, Feldpausch TR, Affum-Baffoe K, Foli EG, Hamer KC, Swaine MD Drought-induced shifts in the floristic and functional composition of tropical forests in Ghana ECOLOGY LETTERS 15 1120-1129, 2012

Riou S, Chastel O, Hamer KC Parent-offspring conflict during the transition to independence in a pelagic seabird Behavioral Ecology 23 1102-1107, 2012
View abstract

Edwards DP, Backhouse AR, Wheeler C, Khen CV, Hamer KC Impacts of logging and rehabilitation on invertebrate communities in tropical rainforests of northern Borneo JOURNAL OF INSECT CONSERVATION 16 591-599, 2012

Woodcock P, Edwards DP, Newton RJ, Edwards FA, Khen CV, Bottrell SH, Hamer KC Assessing trophic position from nitrogen isotope ratios: effective calibration against spatially varying baselines. Naturwissenschaften 99 275-283, 2012
View abstract

Lee Cruz L, McGill RAR, Goodman SJ, Hamer KC Stable isotope ratios of a tropical marine predator: confounding effects of nutritional status during growth Marine Biology 1-8, 2012

Grecian WJ, Witt MJ, Attrill MJ, Bearhop S, Godley BJ, Gremillet D, Hamer KC, Votier SC A novel projection technique to identify important at-sea areas for seabird conservation: An example using Northern gannets breeding in the North East Atlantic BIOLOGICAL CONSERVATION 156 43-52, 2012

Fauset S, Baker TR, Lewis SL, Feldpausch TR, Affum-Baffoe K, Foli EG, Hamer KC, Swaine MD Drought-induced shifts in the floristic and functional composition of tropical forests in Ghana Ecology Letters 15 1120-1129, 2012
View abstract

Woodcock P, Edwards DP, Edwards FA, Hamer KC, Newton RJ, Bottrell SH, Khen CV, Edwards DP Assessing trophic position from nitrogen isotope ratios: effective calibration against spatially varying baselines Naturwissenschaften 1-9, 2012
View abstract

Hill JK, Gray MA, Khen CV, Benedick S, Tawatao N, Hamer KC Ecological impacts of tropical forest fragmentation: how consistent are patterns in species richness and nestedness? PHILOS T R SOC B 366 3265-3276, 2011

Woodcock P, Edwards DP, Fayle TM, Newton RJ, Khen CV, Bottrell SH, Hamer KC The conservation value of South East Asia's highly degraded forests: evidence from leaf-litter ants. Philos Trans R Soc Lond B Biol Sci 366 3256-3264, 2011
View abstract

Xavier JC, Magalhães MC, Mendonç AS, Antunes M, Carvalho N, Machete M, Santos RS, Paiva V, Hamer KC Changes in diet of cory's shearwaters calonectris diomedea breeding in the azores Marine Ornithology 39 129-134, 2011
View abstract

Harding AMA, Welcker J, Steen H, Hamer KC, Kitaysky AS, Fort J, Talbot SL, Cornick LA, Karnovsky NJ, Gabrielsen GW, Gremillet D Adverse foraging conditions may impact body mass and survival of a high Arctic seabird OECOLOGIA 167 49-59, 2011

Ansell FA, Edwards DP, Hamer KC Rehabilitation of Logged Rain Forests: Avifaunal Composition, Habitat Structure, and Implications for Biodiversity-Friendly REDD+ BIOTROPICA 43 504-511, 2011

Bensusan KJ, Shorrocks B, Hamer KC Impacts of passage migrant songbirds on behaviour and habitat use of resident Sardinian Warblers Sylvia melanocephala in Gibraltar IBIS 153 616-621, 2011

Oswald SA, Huntley B, Collingham YC, Russell DJF, Anderson BJ, Arnold JM, Furness RW, Hamer KC Physiological effects of climate on distributions of endothermic species J BIOGEOGR 38 430-438, 2011

Edwards DP, Larsen TH, Docherty TDS, Ansell FA, Hsu WW, Derhe MA, Hamer KC, Wilcove DS Degraded lands worth protecting: the biological importance of Southeast Asia's repeatedly logged forests P ROY SOC B-BIOL SCI 278 82-90, 2011

Riou S, Gray CM, Brooke MD, Quillfeldt P, Masello JF, Perrins C, Hamer KC Recent impacts of anthropogenic climate change on a higher marine predator in western Britain MAR ECOL-PROG SER 422 105-112, 2011

Sodhi NS, Koh LP, Clements R, Wanger TC, Hill JK, Hamer KC, Clough Y, Tscharntke T, Posa MRC, Lee TM Conserving Southeast Asian forest biodiversity in human-modified landscapes BIOL CONSERV 143 2375-2384, 2010

Dunn JC, Hamer KC, Benton TG Fear for the family has negative consequences: indirect effects of nest predators on chick growth in a farmland bird J APPL ECOL 47 994-1002, 2010

Ashbrook K, Wanless S, Harris MP, Hamer KC Impacts of poor food availability on positive density dependence in a highly colonial seabird Proceedings. Biological sciences / The Royal Society 277 2355-2360, 2010
View abstract

Ashbrook K, Wanless S, Harris MP, Hamer KC Impacts of poor food availability on positive density dependence in a highly colonial seabird P ROY SOC B-BIOL SCI 277 2355-2360, 2010

Riou S, Chastel O, Lacroix A, Hamer KC Stress and parental care: Prolactin responses to acute stress throughout the breeding cycle in a long-lived bird GEN COMP ENDOCR 168 8-13, 2010

Edwards DP, Hodgson JA, Hamer KC, Mitchell SL, Ahmad AH, Cornell SJ, Wilcove DS Wildlife-friendly oil palm plantations fail to protect biodiversity effectively CONSERV LETT 3 236-242, 2010

Berry NJ, Phillips OL, Lewis SL, Hill JK, Edwards DP, Tawatao NB, Ahmad N, Magintan D, Khen CV, Maryati M, Ong RC, Hamer KC The high value of logged tropical forests: lessons from northern Borneo BIODIVERS CONSERV 19 985-997, 2010

Riou S, Hamer KC Lipid metabolism, begging behaviour and nestling obesity in a pelagic seabird FUNCT ECOL 24 340-346, 2010

Thaxter CB, Wanless S, Daunt F, Harris MP, Benvenuti S, Watanuki Y, Gremillet D, Hamer KC Influence of wing loading on the trade-off between pursuit-diving and flight in common guillemots and razorbills J EXP BIOL 213 1018-1025, 2010

Dunn JC, Hamer KC, Benton TG Nest and foraging-site selection in Yellowhammers Emberiza citrinella: implications for chick provisioning BIRD STUDY 57 531-539, 2010

Hamer KC The search for winners and losers in a sea of climate change IBIS 152 3-5, 2010

Edwards DP, Ansell FA, Woodcock P, Fayle TM, Chey VK, Hamer KC Can the failure to punish promote cheating in mutualism? OIKOS 119 45-52, 2010

Edwards DP, Ansell FA, Ahmad AH, Nilus R, Hamer KC The Value of Rehabilitating Logged Rainforest for Birds CONSERV BIOL 23 1628-1633, 2009

Harding AMA, Kitaysky AS, Hamer KC, Hall ME, Welcker J, Talbot SL, Karnovsky NJ, Gabrielsen GW, Gremillet D Impacts of experimentally increased foraging effort on the family: offspring sex matters ANIM BEHAV 78 321-328, 2009

Hamer KC, Humphreys EM, Magalhaes MC, Garthe S, Hennicke J, Peters G, Gremillet D, Skov H, Wanless S Fine-scale foraging behaviour of a medium-ranging marine predator J ANIM ECOL 78 880-889, 2009

Harding AMA, Kitaysky AS, Hall ME, Welcker J, Karnovsky NJ, Talbot SL, Hamer KC, Gremillet D Flexibility in the parental effort of an Arctic-breeding seabird FUNCT ECOL 23 348-358, 2009

Thaxter CB, Daunt F, Hamer KC, Watanuki Y, Harris MP, Gremillet D, Peters G, Wanless S Sex-specific food provisioning in a monomorphic seabird, the common guillemot Uria aalge: nest defence, foraging efficiency or parental effort? J AVIAN BIOL 40 75-84, 2009

Ashbrook K, Wanless S, Harris MP, Hamer KC Hitting the buffers: conspecific aggression undermines benefits of colonial breeding under adverse conditions BIOL LETTERS 4 630-633, 2008

Hamer KC Procellariiformes In Encyclopedia of Ocean Sciences: Second Edition, 2008

Riou S, Hamer KC Predation risk and reproductive effort: impacts of moonlight on food provisioning and chick growth in Manx shearwaters ANIM BEHAV 76 1743-1748, 2008

Berry NJ, Phillips OL, Ong RC, Hamer KC Impacts of selective logging on tree diversity across a rainforest landscape: the importance of spatial scale LANDSCAPE ECOL 23 915-929, 2008

Blackall TD, Wilson LJ, Bull J, Theobald MR, Bacon PJ, Hamer KC, Wanless S, Sutton MA Temporal variation in atmospheric ammonia concentrations above seabird colonies ATMOS ENVIRON 42 6942-6950, 2008

Skov H, Humphreys E, Garthe S, Geitner K, Gremillet D, Hamer KC, Hennicke J, Parner H, Wanless S Application of habitat suitability modelling to tracking data of marine animals as a means of analyzing their feeding habitats ECOL MODEL 212 504-512, 2008

Oswald SA, Bearhop S, Furness RW, Huntley B, Hamer KC Heat stress in a high-latitude seabird: effects of temperature and food supply on bathing and nest attendance of great skuas Catharacta skua J AVIAN BIOL 39 163-169, 2008

Daunt F, Wanless S, Greenstreet SPR, Jensen H, Hamer KC, Harris MP The impact of the sandeel fishery closure on seabird food consumption, distribution, and productivity in the northwestern North Sea CAN J FISH AQUAT SCI 65 362-381, 2008

Magalhaes MC, Santos RS, Hamer KC Dual-foraging of Cory's shearwaters in the Azores: feeding locations, behaviour at sea and implications for food provisioning of chicks MAR ECOL-PROG SER 359 283-293, 2008

Benedick S, White TA, Searle JB, Hamer KC, Mustaffa N, Vun Khen C, Mohamed M, Schilthuizen M, Hill JK Impacts of habitat fragmentation on genetic diversity in a tropical forest butterfly on Borneo J TROP ECOL 23 623-634, 2007

Blackall TD, Wilson LJ, Theobald MR, Milford C, Nemitz E, Bull J, Bacon PJ, Hamer KC, Wanless S, Sutton MA Ammonia emissions from seabird colonies Geophysical Research Letters 34 pp.L10801-, 2007
View abstract

Hamer KC, Humphreys EM, Garthe S, Hennicke J, Peters G, Gremillet D, Phillips RA, Harris MP, Wanless S Annual variation in diets, feeding locations and foraging behaviour of gannets in the North Sea: flexibility, consistency and constraint MAR ECOL-PROG SER 338 295-305, 2007

Benedick S, Hill JK, Mustaffa N, Chey VK, Maryati M, Searle JB, Schilthuizen M, Hamer KC Impacts of rain forest fragmentation on butterflies in northern Borneo: species richness, turnover and the value of small fragments J APPL ECOL 43 967-977, 2006

Hamer KC, Lewis S, Wanless S, Phillips RA, Sherratt TN, Humphreys EM, Hennicke J, Garthe S Use of gannets to monitor prey availability in the northeast Atlantic Ocean: colony size, diet and foraging behaviour TOP PREDATORS IN MARINE ECOSYSTEMS: THEIR ROLE IN MONITORING AND MANAGEMENT 236-248, 2006

Enstipp MR, Daunt F, Wanless S, Humphreys EM, Hamer KC, Benvenuti S, Gremillet D Foraging energetics of North Sea birds confronted with fluctuating prey availability TOP PREDATORS IN MARINE ECOSYSTEMS: THEIR ROLE IN MONITORING AND MANAGEMENT 191-210, 2006

Hamer KC, Hill JK, Benedick S, Mustaffa N, Chey VK, Maryati M Diversity and ecology of carrion- and fruit-feeding butterflies in Bornean rain forest Journal of Tropical Ecology 22 25-33, 2006
View abstract

Hamer KC, Quillfeldt P, Masello JF, Fletcher KL Sex differences in provisioning rules: responses of Manx shearwaters to supplementary chick feeding BEHAV ECOL 17 132-137, 2006

Gray CM, Brooke MDL, Hamer KC Repeatability of chick growth and food provisioning in Manx shearwaters Puffinus puffinus Journal of Avian Biology 36 374-379, 2005
View abstract

Hamer KC, Hill JK, Mustaffa N, Benedick S, Sherratt TN, Chey VK, Maryati M Temporal variation in abundance and diversity of butterflies in Bornean rain forests: opposite impacts of logging recorded in different seasons Journal of Tropical Ecology 21 417-425, 2005
View abstract

Lewis S, Schreiber EA, Daunt F, Schenk GA, Orr K, Adams A, Wanless S, Hamer KC Sex-specific foraging behaviour in tropical boobies: does size matter? IBIS 147 408-414, 2005

Lewis S, Schreiber EA, Daunt F, Schenk GA, Wanless S, Hamer KC Flexible foraging patterns under different time constraints in tropical boobies ANIM BEHAV 68 1331-1337, 2004

Wilson LJ, Bacon PJ, Bull J, Dragosits U, Blackall TD, Dunn TE, Hamer KC, Sutton MA, Wanless S Modelling the spatial distribution of ammonia emissions from seabirds in the UK Environmental Pollution 131 173-185, 2004
View abstract

Quillfeldt P, Masello JF, Hamer KC Sex differences in provisioning rules and honest signalling of need in Manx shearwaters, Puffinus puffinus ANIM BEHAV 68 613-620, 2004

Hill JK, Hamer KC Determining impacts of habitat modification on diversity of tropical forest fauna: the importance of spatial scale J APPL ECOL 41 744-754, 2004

Fletcher KL, Hamer KC Offspring sex ratio in the Common Tern Sterna hirundo, a species with negligible sexual size dimorphism Ibis 146 454-460, 2004
View abstract

Lewis S, Hamer KC, Money L, Griffiths R, Wanless S, Sherratt TN Brood neglect and contingent foraging behavior in a pelagic seabird BEHAV ECOL SOCIOBIOL 56 81-88, 2004

Votier SC, Furness RW, Bearhop S, Crane JE, Caldow RWG, Catry P, Ensor K, Hamer KC, Hudson AV, Kalmbach E, Klomp NI, Pfeiffer S, Phillips RA, Prieto I, Thompson DR Changes in fisheries discard rates and seabird communities NATURE 427 727-730, 2004

Wilson LJ, Bacon PJ, Bull J, Dragosits U, McDonald A, Blackall TD, Dunn T, Hamer KC, Sutton MA, Wanless S The spatial distribution of ammonia emitted from seabirds and its contribution to atmospheric nitrogen deposition in the UK Water, Air and Soil Pollution 4 287-296, 2004

Lewis S, Benvenuti S, Daunt F, Wanless S, Dall'Antonia L, Luschi P, Elston DA, Hamer KC, Sherratt TN Partitioning of diving effort in foraging trips of northern gannets Canadian Journal of Zoology 82 1910-1916, 2004

Hay JM, Evans PR, Ward RM, Hamer KC Poor nutritional condition as a consequence of high dominance status in the Coal Tit Parus ater Ibis 146 103-107, 2004

Blackall TD, Theobald MR, Milford C, Hargreaves KJ, Nemitz E, Wilson LJ, Bull J, Bacon PJ, Hamer KC, Wanless S, Sutton MA Application of tracer ratio and inverse dispersion methods with boat-based measurements to estimate ammonia emissions fom seabird colonies Water, Air and Soil Pollution 4 279-285, 2004

Hamer KC Puffinus puffinus Manx shearwater BWP Update 5 203-213, 2003
View abstract

Fletcher KL, Hamer KC Sexing terns using biometrics: the advantage of within-pair comparisons Bird Study 50 78-83, 2003

Hamer KC, Hill JK, Benedick S, Mustaffa N, Sherratt TN, Maryati M, Chey VK Ecology of butterflies in natural and selectively logged forests of northern Borneo: the importance of habitat heterogeneity Journal of Applied Ecology 40 150-162, 2003

Gray CM, Phillips RA, Hamer KC Non-random mortality of nestling fulmars, Fulmarus glacialis: implications for monitoring marine environments Journal of Zoology (London, 1987) 259 109-114, 2003

Lewis S, Sherratt TN, Hamer KC, Harris MP, Wanless S Contrasting diet quality of northern gannets Morus bassanus at two colonies ARDEA 91 167-176, 2003

Hill JK, Hamer KC, Dawood M, Tangah J, Chey VK Rainfall but not selective logging affects changes in abundance of a tropical forest butterfly in Sabah, Borneo Journal of Tropical Ecology 19 35-42, 2003

Harding AMA, Piatt JF, Hamer KC Breeding ecology of horned puffins (Fratercula corniculata) in Alaska: annual variation and effects of El Nino Canadian Journal of Zoology 81 1004-1013, 2003

Lewis S, Benvenuti S, Dall'Antonia L, Griffiths R, Money L, Sherratt TN, Wanless S, Hamer KC Sex-specific foraging behaviour in a monomorphic seabird Proceedings of the Royal Society of London. Series B. Biological Sciences 269 1687-1693, 2002

Ratcliffe N, Catry P, Hamer KC, Klomp NI, Furness RW The effect of age and year on the survival of breeding adult Great Skuas Catharacta skua in Shetland Ibis 144 384-392, 2002

Robinson JA, Hamer KC, Chivers LS Developmental plasticity in Arctic Terns Sterna paradisaea and Common Terns S-hirundo in response to a period of extremely bad weather Ibis 144 344-346, 2002

Redman KK, Lewis S, Griffiths R, Wanless S, Hamer KC Sexing northern gannets from DNA, morphology and behaviour Waterbirds 25 230-234, 2002

Bearhop S, Thompson DR, Phillips RA, Waldron S, Hamer KC, Gray CM, Votier SC, Ross BP, Furness RW Annual variation in Great Skua diets: The importance of commercial fisheries and predation on seabirds revealed by combining dietary analyses Condor (Tempe) 103 802-809, 2001

Gray CM, Hamer KC Prefledgling mass recession in Manx shearwaters: parental desertion or nestling anorexia? Animal Behaviour 62 705-709, 2001

Hamer KC Nestling obesity in shearwaters: a reply to Gaston (2001) Ibis 143 686-687, 2001

Lewis S, Sherratt TN, Hamer KC, Wanless S Evidence of intra-specific competition for food in a pelagic seabird Nature 412 816-819, 2001

Hill JK, Hamer KC, Tangah J, Dawood M Ecology of tropical butterflies in rainforest gaps Oecologia 128 294-302, 2001

Gray CM, Hamer KC Food-provisioning behaviour of male and female Manx shearwaters, Puffinus puffinus Animal Behaviour 62 117-121, 2001

Robinson JA, Hamer KC, Chivers LS Contrasting brood sizes in Common and Arctic Terns: The roles of food provisioning rates and parental brooding Condor (Tempe) 103 108-117, 2001

Hamer KC Great Skua BWP Update 3 91-110, 2001
View abstract

Hamer KC, Phillips RA, Hill JK, Wanless S, Wood AG Contrasting foraging strategies of gannets Morus bassanus at two North Atlantic colonies: foraging trip duration and foraging area fidelity Marine Ecology - Progress Series 224 283-290, 2001

Bradley JS, Cox JM, Nicholson LW, Wooller RD, Hamer KC, Hill JK Parental influence upon the provisioning schedules of nestling Short-tailed Shearwaters Puffinus tenuirostris J AVIAN BIOL 31 522-526, 2000

Hamer KC, Hill JK Scale-dependent effects of habitat disturbance on species richness in tropical forests CONSERV BIOL 14 1435-1440, 2000

Phillips RA, Hamer KC Postnatal development of northern fulmar chicks, Fulmarus glacialis. Physiol Biochem Zool 73 597-604, 2000
View abstract

Phillips RA, Hamer KC Growth and provisioning strategies of Northern Fulmars Fulmarus glacialis IBIS 142 435-445, 2000

Phillips RA, Hamer KC Periodic weighing and the assessment of meal mass and feeding frequency in seabirds J AVIAN BIOL 31 75-80, 2000

Thompson DR, Hamer KC Stress in seabirds: Causes, consequences and diagnostic value Journal of Aquatic Ecosystem Stress and Recovery 7 91-110, 2000
View abstract

Hamer KC, Phillips RA, Wanless S, Harris MP, Wood AG Foraging ranges, diets and feeding locations of gannets Morus bassanus in the North Sea: evidence from satellite telemetry MAR ECOL-PROG SER 200 257-264, 2000

Robinson JA, Hamer KC Brood size and food provisioning in common terns Sterna hirundo and arctic terns S-paradisaea: Consequences for chick growth ARDEA 88 51-60, 2000

Hamer KC, Hill JK, Bradley JS, Wooller RD Contrasting patterns of nestling obesity and food provisioning in three species of Puffinus shearwaters: the role of predictability IBIS 142 146-150, 2000

Phillips RA, Petersen MK, Lilliendahl K, Solmundsson J, Hamer KC, Camphuysen CJ, Zonfrillo B Diet of the northern fulmar Fulmarus glacialis: reliance on commercial fisheries? MAR BIOL 135 159-170, 1999

Hamer KC, Hill JK, Scott I Chick provisioning and parental attendance in Cory's Shearwaters: implications for nestling obesity J AVIAN BIOL 30 309-315, 1999

Phillips RA, Hamer KC Lipid reserves, fasting capability and the evolution of nestling obesity in procellariiform seabirds P ROY SOC LOND B BIO 266 1329-1334, 1999

Phillips RA, Bearhop S, Hamer KC, Thompson DR Rapid population growth of Great Skuas Catharacta skua at St Kilda: implications for management and conservation BIRD STUDY 46 174-183, 1999

Phillips RA, Thompson DR, Hamer KC The impact of great skua predation on seabird populations at St Kilda: a bioenergetics model J APPL ECOL 36 218-232, 1999

Hamer KC, Lynnes AS, Hill JK Parent-offspring interactions in food provisioning of Manx shearwaters: implications for nestling obesity. Anim Behav 57 627-631, 1999
View abstract

Catry P, Ruxton GD, Ratcliffe N, Hamer KC, Furness RW Short-lived repeatabilities in long-lived great skuas: implications for the study of individual quality OIKOS 84 473-479, 1999

Ratcliffe N, Furness RW, Hamer KC The interactive effects of age and food supply on the breeding ecology of great skuas J ANIM ECOL 67 853-862, 1998

Royle NJ, Hamer KC Hatching asynchrony and sibling size hierarchies in gulls: effects on parental investment decisions, brood reduction and reproductive success J AVIAN BIOL 29 266-272, 1998

Hamer KC, Lynnes AS, Hill JK Regulation of chick provisioning rate in Manx Shearwaters: experimental evidence and implications for nestling obesity FUNCT ECOL 12 625-630, 1998

Catry P, Phillips RA, Hamer KC, Ratcliffe N, Furness RW The incidence of nonbreeding by adult Great Skuas and Parasitic Jaegers from Foula, Shetland CONDOR 100 448-455, 1998

Hill JK, Hamer KC, Hodkinson ID Variation in resource exploitation along an altitudinal gradient: the willow psyllids (Cacopsylla spp.) on Salix lapponum ECOGRAPHY 21 289-296, 1998

Hill JK, Hamer KC Using species abundance models as indicators of habitat disturbance in tropical forests J APPL ECOL 35 458-460, 1998

Hamer KC, Hill JK Nestling obesity and variability of food delivery in Manx Shearwaters, Puffinus puffinus FUNCT ECOL 11 489-497, 1997

Cook MI, Hamer KC Effects of supplementary feeding on provisioning and growth rates of nestling Puffins Fratercula arctica: Evidence for regulation of growth J AVIAN BIOL 28 56-62, 1997

Hamer KC, HIll JK, Lace LA, Langan AM Ecological and biogeographical effects of forest disturbance on tropical butterflies of Sumba, Indonesia J BIOGEOGR 24 67-75, 1997

Phillips RA, Catry P, Thompson DR, Hamer KC, Furness RW Inter-colony variation in diet and reproductive performance of great skuas Catharacta skua MAR ECOL-PROG SER 152 285-293, 1997

Hamer KC, Nicholson LW, Hill JK, Wooller RD, Bradley JS Nestling obesity in procellariiform seabirds: temporal and stochastic variation in provisioning and growth of short-tailed shearwaters Puffinus tenuirostris OECOLOGIA 112 4-11, 1997

Hamer KC, Thompson DR Provisioning and growth rates of nestling Fulmars Fulmarus glacialis: Stochastic variation or regulation? IBIS 139 31-39, 1997

Hill JK, Hamer KC, Lace LA, Banham WMT Effects of selective logging on tropical forest butterflies on Buru, Indonesia. Journal of Applied Ecology 32 754-760, 1995
View abstract








Hamer KC, Furness RW Sexing great skuas catharacta skua by discriminant analysis using external measurements Ringing and Migration 12 16-22, 1991
View abstract

Jones MJ, Lace LA, Hounsome MV, Hamer K The birds and butterflies of Madeira and La Gomera: taxon cycles and human influence Biological Journal of the Linnean Society 31 95-111, 1987